The Village of Speculator is the economic and social center of the town with many dining, lodging, and shopping facilities. Incorporated in 1925, the village is a center for activities such as downhill skiing, snowmobiling, mountain biking, swimming, tennis, basketball, and much more. Speculator gained world-wide notoriety shortly after World War I, when it became the preferred training site for such heavy-weight boxing champions as Gene Tunney, Max Baer, and Max Schmelling.

For those interested in the history and environment of the area, Speculator boasts an interpretive river walk called Sacandaga River Community Park. This handicapped-accessible pathway is visited and enjoyed by thousands of people each year; it consists of boardwalks and trails starting at the ball field and winding through an Adirondack wetland.